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2007. Embryology of Swertia (Gentianaceae) relative to taxonomy. BOJ Botanical Journal of the Linnean Society. 155(3):383-400.
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2005. Embryology of Megacodon stylophorus and Veratrilla baillonii (Gentianaceae): descriptions and systematic implications. BOJ Botanical Journal of the Linnean Society. 147(3):317-331.